Tuula Hollmen

Tuula Hollmen

Research Associate Professor

Alaska Sealife Center
PO Box 1329
Seward, AK 99664



University of Helsinki

University of Helsinki


Curriculum Vitae




Ellsworth, A., Buck, C.L., Atkinson, S., Hollmen, T.. (2014). "Longitudinal monitoring of sex steroid hormones in excrement of spectacled eiders (Somateria fisheri)". Gen. Comp. Endocrinol.. 189:59-65.

C. Frost, T. Hollmen and J. Reynolds. (2013). "Trends in annual survival of Steller's eiders molting at Izembek Lagoon on the Alaska Peninsula 1993-2006".  Arctic. 66:173-178.

M. W. Seltmann, M. Ost, K. Jaatinen, S. Atkinson, K. Mashburn and T. Hollmen. (2012). "Stress responsiveness, age and body condition interactively affect flight initiation distance in breeding female eiders". Animal Behaviour. 84(4):889-896.

R. N. Federer, T. E. Hollmen, D. Esler and M. J. Wooller. (2012). "Stable Carbon and Nitrogen Isotope Discrimination Factors for Quantifying Spectacled Eider Nutrient Allocation to Egg Production Condor". 114(4):726-732.

S. M. Budge, S. W. Wang, T. E. Hollmen and M. J. Wooller. (2011). "Carbon isotopic fractionation in eider adipose tissue varies with fatty acid structure: implications for trophic studies". Journal of Experimental Biology. 214(22):3790-3800.

T. E. Hollmen, C. DebRoy, P. L. Flint, D. E. Safine, J. L. Schamber, A. E. Riddle and K. A. Trust. (2011). "Molecular typing of Escherichia coli strains associated with threatened sea ducks and near-shore marine habitats of south-west Alaska". Environmental Microbiology Reports. 3(2):262-269.

C. J. Latty, T. E. Hollmen, M. R. Petersen, A. N. Powell and R. D. Andrews. (2010). "Abdominally implanted transmitters with percutaneous antennas affect the dive performance of common eiders". Condor. 112(2):314-322.
doi: 10.1525/cond.2010.090022

R. N. Federer, T. E. Hollmen, D. Esler, M. J. Wooller and S. W. Wang. (2010). "Stable carbon and nitrogen isotope discrimination factors from diet to blood plasma, cellular blood, feathers, and adipose tissue fatty acids in Spectacled Eiders (Somateria fischeri)". Canadian Journal of Zoology-Revue Canadienne De Zoologie. 88(9):866-874.
doi: 10.1139/z10-052

S. Oppel, R. N. Federer, D. M. O'Brien, A. N. Powell and T. E. Hollmen. (2010). "Effects of lipid extraction on stable isotope ratios in avian egg yolk: is arithmetic correction a reliable alternative?". Auk. 127(1):72-78.
doi: 10.1525/auk.2009.09153

S. W. Wang, T. E. Hollmen and S. J. Iverson. (2010). "Validating quantitative fatty acid signature analysis to estimate diets of spectacled and Steller's eiders (Somateria fischeri and Polysticta stelleri)". Journal of Comparative Physiology B-Biochemical Systemic and Environmental Physiology. 180(1):125-139.
doi: 10.1007/s00360-009-0393-x


  • Physiology and health of wild avian species, especially waterfowl and marine birds
  • Development of methods and biomarkers to evaluate body condition, immune function, and disease/contaminant effects in birds
  • Application of such techniques for monitoring of population health and fitness of wild birds